Dueling puffins flare their wings and part their beaks in a muddy tussle on Skomer Island, Wales. Most puffin fights start over competing claims to a burrow. A bite to the neck usually settles the issue.
Most puffins start breeding when they are five years old and often live for more than 20 years. Some young, inexperienced birds may change mates after breeding failures but most will mate with the same partner for many years.